Barr, D. J. 2004 Establishing conventional communication systems: is common knowledge necessary? Cognit. Sci. 28, 937-962. (doi:10.1016/j.cogsci.2004.07.002)

Basalla, G. 1988 The evolution of technology. Cambridge, UK: Cambridge University Press.

Caldwell, C. A. & Millen, A. E. 2008 Experimental models for testing hypotheses about cumulative cultural evolution. Evol. Hum. Behav., 165-171. (doi:10.1016/ j.evolhumbehav.2007.12.001)

Caldwell, C. A. & Millen, A. E. 2008 Studying cumulative cultural evolution in the laboratory. Phil. Trans. R. Soc. B 363, 3529-3539. (doi:10.1098/rstb.2008.0133)

Christiansen, M. H. & Kirby, S. 2003 Language evolution: consensus and controversies. Trends Cognit. Sci. 7, 300-307. (doi:10.1016/S1364-6613(03)00136-0)

Clark, H. H. 1996 Using language. Cambridge, UK: Cambridge University Press.

Dediu, D. D. & Ladd, D. R. D. R. 2007 Linguistic tone is related to the population frequency of the adaptive haplogroups of two brain size genes, ASPM and Microcephalin. Proc. Natl Acad. Sci. USA 104, 10 944-10 949. (doi:10.1073/pnas.0610848104)

Fay, N., Garrod, S., Roberts, L. & Swoboda, N. In preparation. The interactive evolution of communication systems.

Galantucci, B. 2005 An experimental study of the emergence of human communication systems. Cognit. Sci. 29, 737-767. (doi:10.1207/s15516709cog0000_34)

Garrod, S. & Doherty, G. 1994 Conversation, coordination and convention--an empirical investigation of how groups establish linguistic conventions. Cognition 53, 181-215. (doi:10.1016/0010-0277(94)90048-5)

Garrod, S., Fay, N., Lee, J., Oberlander, J. & MacLeod, T. 2007 Foundations of representation: where might graphical symbol systems come from? Cognit. Sci. 31, 961-987. (doi:10.1080/03640210701703659)

Gittins, D. 1986 Icon-based human computer interaction. Int. J. Man-Machine Stud. 24, 519-543.

Goldin-Meadow, S. 2003 The resilience of language: what gesture creation in deaf children can tell us about how children learn language. New York, NY: Psychology Press.

Healy, G. T., Swoboda, N. & King, J. 2002 A tool for performing and analysing experiments on graphical communication. In People and computers XVI: Proc. HCI2002: The 16th British HCI Group Annual Conference (eds X. Faulkner, J. Finlay & F. Detienne), pp. 55-68. London, UK: Springer.

Healy, P. G. T., Swoboda, N., Umata, I. & King, J. 2007 Graphical language games: interactional constraints on representational form. Cognit. Sci. 31, 285-309. (doi:10. 1080/15326900701221363)

Kegl, J., Senghas, A. & Coppola, M. 1999 Creation through contact: sign language emergence and sign language change in Nicaragua. In Language creation and language change: creolization, diachrony and development (ed. M. DeGraff), pp. 179-237. Cambridge, MA: MIT Press.

Kirby, S. 2002 Natural language from artificial life. Artif. Life 8, 185-215. (doi:10.1162/106454602320184248)

Kirby, S. & Hurford, J. 1997 Learning, culture and evolution in the origin of linguistic constraints. In Proc. Fourth European Conference on Artificial Life (eds S. Husbands & I. Harvey), pp. 493-502. Cambridge, MA: MIT Press.

Kirby, S. & Hurford, J. 2002 The emergence of linguistic structure: an overview of the iterated learning model. In Simulating the evolution of language (eds A. Cangelosi & D. Parisi), pp. 121-148. London, UK: Springer.

Kirby, S., Dowman, M. & Griffiths, T. L. 2007 Innateness and culture in the evolution of language. Proc. Natl Acad. Sci. USA 104, 5241-5245. (doi:10.1073/pnas.0608222104)

Lieberman, E., Michel, J. B., Jackson, J., Tang, T. & Nowak, M. A. 2007 Quantifying the evolutionary dynamics of language.Nature 449, 713-716. (doi:10.1038/nature06137)

McDougall, S. 2000 Exploring the effects of icon characteristics on user performance: the role of icon concreteness, complexity, and distinctiveness. J. Exp. Psychol. Appl. 6, 291-306. (doi:10.1037/1076-898X.6.4.291)

Mesoudi, A., Whiten, A. & Laland, K. N. 2006 Towards a unified science of cultural evolution. Behav. Brain Sci. 29, 329-347. (doi:10.1017/S0140525X06009083)

Pagel, M., Atkinson, Q. D. & Meade, A. 2007 Frequency of word-use predicts rates of lexical evolution throughout Indo-European history. Nature 449, 717-U717. (doi:10. 1038/nature06176)

Pickering, M. J. & Garrod, S. 2004 Toward a mechanistic psychology of dialogue. Behav. Brain Sci. 27, 169-190. (doi:10.1017/S0140525X04000056)

Pinker, S. 1994 The language instinct. New York, NY: HarperCollins.

Pinker, S. & Jackendoff, R. 2005 The faculty of language: what's special about it? Cognition 95, 201-236. (doi:10. 1016/j.cognition.2004.08.004)

Plotkin, H. C. 2002 The imagined world made real. Harmondsworth, UK: Penguin.

Sandler, W., Meir, I., Padden, C. & Aronoff, M. 2005 The emergence of grammar: systematic structure in a new language. Proc. Natl Acad. Sci. USA 102, 2661-2665. (doi:10.1073/pnas.0405448102)

Seyfarth, R. M. & Cheney, D. L. 2003 Signalers and receivers in animal communication. Annu. Rev. Psychol. 54, 145-173. (doi:10.1146/annurev.psych.54.101601.145121)

Steels, L., Kaplan, F., McIntyre, A. & Van Looveren, J. 2002 Crucial factors in the origins of word-meaning. In The transition to language (ed. A. Wray), pp. 252-271. Oxford, UK: Oxford University Press.

Taylor, M. M. & Creelman, C. D. 1967 PEST: efficient estimates in probability functions. J. Acoust. Soc. Am. 4, 782-787. (doi:10.1121/1.1910407)

Tomasello, M. 1999 The cultural origins of human cognition. Cambridge, MA: Harvard University Press.

Tomasello, M., Carpenter, M., Call, J., Behne, T. & Moll, H. 2005 Understanding and sharing intentions: the origins of cultural cognition. Behav. Brain Sci. 28, 675-691. (doi:10. 1017/S0140525X05000129)

Treutwein, B. 1995 Adaptive psychophysical procedures. Vis. Res. 35, 2503-2522. (doi:10.1016/0042-6989(95) 00016-X) Culturally evolved communication systems N. Fay et al. 3561